The aim of this study was to investigate relationships between soil properties and plant species to determine the most effective factors separating vegetation communities in Rineh rangeland. Three stratifying variables were selected including slop, aspect and elevation. The study area was partitioned by combining these classes to generate homogenous units. ۱m۲ quadrates were located at sampling sites in each homogenous unit randomly.
Vegetation cover data were recorded using ordinal scale of Daubenmire cover-abundance scores in each quadrate. Soil samples were collected based on area from ۰-۱۵, ۱۵-۳۰ cm depths in each sampling site. Total nitrogen, organic matter, pH, total phosphorus, water retention capacity, permanent wilting point, available water capacity, potassium, water holding capacity, CaCO۳, saturation moisture, bulk density, real unit weight and percentage of fine earth fragments (sand, silt and clay) were measured. Both classification and ordination techniques were employed including TWINSPAN classification, DCA and CCA. The TWINSPAN classification of the sample sites have resulted in ten groups. According to the results of DCA, length of gradient represented by axis ۱ was >۵ SD, indicating that CCA was the appropriate ordination method. CCA axis ۱ was correlated to phosphorus (-۰.۴۶۰) in the first layer and phosphorus (-۰.۴۹۳), sand (۰.۵۳۳) in the second layer while the CCA axis ۲ was correlated to phosphorus (۰.۳۹۴), sand (۰.۵۳۳) in the second layer. The species-environment correlations are higher for the first three canonical axes, explaining ۱۸% of the cumulative variances. REFERENCES Ali, S.M., Malik, N.R. (۲۰۱۰) vegetation communities of urban open species: Green belt and parks in Islamabad city. Pak. J. Bot, ۴۲, ۱۰۳۱-۱۰۳۹. Bouyoucos, G.J. (۱۹۶۲) Hydrometer method improved for making particle size analyses of soils. Argon. J. ۵۴, ۴۶۴-۴۶۵. El Bana, M.I., A.S. Al-Mathnani., (۲۰۰۹) Vegetation-soil relationships in the Wadi Al-Hayat Area of the Libyan Sahara, Aust. J. Basic & Appl. Sci. ۳, ۷۴۰-۷۴۷. El Bana, M.I., A.A. Khedr, P. Van Hecke, and J. Bogaert, (۲۰۰۲) Vegetation composition of a threatened hypersaline, Ecol.۱۶۳, ۶۳-۷۵. El Ghani, M.M. (۱۹۹۸) Environmental correlates of species distribution in arid desert ecosystems of eastern Egypt. J. Arid Environ. ۳۸, ۲۹۷-۳۱۳. He, M.Z., Zheng, J.G., Li, X.R., Qian, Y.L. (۲۰۰۷) Environmental factors affecting vegetation composition in the Alxa Plateau, China. J. Arid Environ. ۶۹, ۴۷۳-۴۸۹. Hirzel, A., and Guisan, A. (۲۰۰۲). Which is optimal sampling strategy for habitat suitability modeling? Ecol. Model. ۱۵۷, ۳۳۱-۳۴۱. Jafari, M., Zare Chahouki, M.A., Tavili, A., Azarnivand, H., Zahedi Amiri, Gh. (۲۰۰۴). Effective environmental factors in the distribution of vegetation types in Poshtkouh rangelands of Yazd Province (Iran). J. Arid Environ. ۵۶, ۶۲۷-۶۴۱. Jafarian Z., H. Arzani, M. Jafari, Gh. Zahedi, H. Azarnivand. (۲۰۰۸) Analyzing the relationship between distribution or plant communities and climatic and physiographic factors using Classification and Ordination methods in Rineh Rangelands. Range. J. ۲, ۱۲۵-۱۴۰. Kabir, M., Zafariqbal, M., Farooqi, Z.R., Shafiq, M. (۲۰۱۰) vegetation pattern and soil characteristics of the polluted industrial area of Karachi. Pak. J. Bot. ۴۲, ۶۶۱-۶۷۸. Kumar, S. (۱۹۹۶). Trends in structural compositional attributes of duneinterdune vegetation and their edaphic relations in the Indian desert. Vegetatio, ۱۲۴, ۷۳-۹۳. Li, X.R., Liu, X.M., Yang, Z.Y. (۱۹۹۸). A study on the relation of shrub community and environment in decertified steppe and steppefied desert of Ordos plateau. Chinese J. of Arid Res., ۱۸, ۱۲۳-۱۳۰. Lu, T., Ma, K.M., Zhang, W.H., Fu, B.J. (۲۰۰۶). Differential responses of shrubs and herbs present at the Upper Minjiang River basin (Tibetan Plateau) to several soil variables. J. Arid Environ. ۶۷, ۳۷۳-۳۹۰. Lundholm, J.T., Larson, D.W. (۲۰۰۳) Relationships between spatial environmental heterogeneity and plant species diversity on a limestone pavement. Ecography. ۲۶, ۷۱۵-۷۲۲. McCune, B., Mefford, M.J. (۱۹۹۹). PC-ORD ۴: Multivariate Analysis of Ecological Data. Version ۴.۱۰. MjM Software, Gleneden Beach, OR, USA. McDonald, D.J., Cowling, R.M., Boucher, C. (۱۹۹۶). Vegetation-environment relationships on a species-rich coastal mountain range in the fynbos biom (South Africa). Vegetatio. ۱۲۳, ۱۶۵-۱۸۲. McGill, W.B., Figueiredo, C.T. (۱۹۹۳) Total nitrogen. In Carter, M.R. Soil Sampling and Methods of Analysis. Lewis Publishers. Boca Raton, FL, pp. ۲۰۱-۲۱۱. Monier, M., El-Ghani Abd., Amwe, W.A. (۲۰۰۳) Soil-vegetation relationships in a costal desert plain of southern Sinai, Egypt. J. Arid Environ. ۵۵, ۶۰۷-۶۲۸. Nelson, D.W., Summers, L.E., et al., Total carbon, organic carbon, and organic matter. In: Page, A.L. (Ed), Methods of Soil Analysis. (۱۹۸۲) Part ۲. Chemical and Microbiological Properties, second ed. Agronomy Monographs, Madison, ۹, ۵۳۹- ۵۷۹. Oksanen, J., Minchin, P.R. (۱۹۹۷) Instability of ordination results under changes in input data order: explanations and remedies. J. Veg. Sci. ۸, ۴۴۷-۴۵۴. Page, A.L., Miller, R.H., Keeney, D.R. (۱۹۸۲) Methods of Soil Analysis. Part ۲. Chemical and Microbiological Properties, second ed. Agronomy Monographs, ASA-SSA, Madison, WI. Palmer, M.W. (۱۹۹۳) Putting thing in an even better order; the advantages of canonical correspondence analysis, Ecology, ۷۴, ۲۲۱۵-۲۲۳۰. Shaukat, S.S., Khan, D., Qadir, S.A. (۱۹۸۱) on the vegetation dynamics of calcareous hillsaround Karachi. Pak. J. Bot. ۱۳, ۱۷-۳۷. Simard, R.R. (۱۹۹۳) Ammonium acetateextractable elements. In: Carter, M.R. (Ed.), Soil Sampling and Methods of Analysis. Lewis Publishers, Boca Rton, FL, pp.۳۹-۴۲. Sperry, J.S., Hacke, U.G. (۲۰۰۲) Desert shrub water relations with respect to soil characteristics and plant functional type. Funct. Ecol. ۱۶, ۳۶۷-۳۷۸. Yair, A., Danin, A. (۱۹۸۰) Spatial variation as related to the soil moisture regime over an arid limestone hillside, northern Negev, Israel. Oncology. ۴۷, ۸۳-۸۸. Zhang, K., Dang, H., Tan, Sh., Wang Zh., Zhang, Q. (۲۰۱۰) Vegetation community and soil characteristics of abandoned agricultural land and pine plantation in the Qinling Mountains, China. Forest Ecol. and Manag. ۲۵۹, ۲۰۳۶-۲۰۴۷.